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Typology: Schemes and Mind Maps
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These are the tissues where active cell division and growth occurs. Based on the position, meristems are 3 types: o Apical meristems: They occur at the tips of roots and shoots and produce primary tissues. It is 2 types: § Root apical meristem: It occupies the tip of a root. § Shoot apical meristem: It occupies the distant most region of the stem axis. Some cells ‘left behind’ from shoot apical meristem, constitute the axillary bud. They are present in the axils of leaves and can form a branch or a flower.
o Intercalary meristems: They occur between mature tissues. They occur in grasses and regenerate parts removed by the grazing herbivores. Apical and intercalary meristems are primary meristems because they appear early in a plant life and contribute to the formation of primary plant body. During that, specific regions of the apical meristem produce dermal tissues, ground tissues and vascular tissues. o Secondary (lateral) meristems: The meristems that occur in mature regions of roots and shoots. They are cylindrical meristems. They are seen in gymnosperms and dicots. E.g. Fascicular vascular cambium, interfascicular cambium & cork cambium. These are responsible for producing the secondary tissues.
a. Parenchyma
o Tracheids: These are elongated tube like dead cells with thick and lignified walls and tapering ends. Protoplasm absent. The inner layers of cell walls have thickenings which vary in form. In flowering plants, tracheids & vessels are the main water transporting elements.
o Vessel: It is a long cylindrical tube-like structure made up of many cells (vessel members), each with lignified walls and a large central cavity. Protoplasm absent. Vessel members are interconnected through perforations in their common walls. The vessels are a characteristic feature of angiosperms. Gymnosperms lack vessels.
o Xylem fibres: They have highly thickened walls and obliterated central lumens. They are septate or aseptate.
o Xylem parenchyma: Living and thin-walled cells with cellulosic cell walls. They store food materials (starch or fat) and other substances like tannins. Radial conduction of water takes place by the ray parenchymatous cells.
o Sieve tube elements: These are long, tube-like structures, arranged longitudinally and are associated with companion cells. Their end walls are perforated to form the sieve plates. A mature sieve element has a peripheral cytoplasm and a large vacuole but lacks a nucleus. The functions of sieve tubes are controlled by the nucleus of companion cells. The first formed primary phloem (protophloem) consists of narrow sieve tubes. The later formed phloem (metaphloem) has bigger sieve tubes. Function: Conduction of food materials from leaves. o Companion cells: Specialized parenchymatous cells closely associated with sieve tube elements. Sieve tube elements & companion cells are connected by pit fields present between their common longitudinal walls. Function: Maintain the pressure gradient in sieve tubes. o Phloem parenchyma: It is made up of elongated, tapering cylindrical cells which have dense cytoplasm and nucleus. The cell wall is composed of cellulose and has pits through which plasmodesmatal connections exist between the cells. Phloem parenchyma is absent in most of the monocots. Function: It stores food material and other substances like resins, latex and mucilage. o Phloem fibres (bast fibres): These are made up of sclerenchymatous cells. Generally absent in primary phloem but are found in the secondary phloem. These are much elongated, unbranched and have pointed, needle like apices. Cell wall is quite thick. At maturity, these fibres lose their protoplasm and become dead. Phloem fibres of jute, flax and hemp are used commercially. Function: Mechanical support & protection to soft tissues.
Based on structure and location, tissue systems are 3 types: o Epidermal tissue system o Ground (fundamental) tissue system o Vascular (conducting) tissue system
Stomata
§ Hypodermis: Outer zone. It consists of a few layers of collenchymatous cells just below the epidermis. It provides mechanical strength to the young stem. § Cortical layers: Below hypodermis. They consist of rounded thin walled parenchymatous cells with conspicuous intercellular spaces. § Endodermis: Innermost layer. The cells are rich in starch grains. So the layer is also called as the starch sheath. Pericycle is present on the inner side of the endodermis and above the phloem in the form of semi- lunar patches of sclerenchyma. o Stele: Consists of pericycle, vascular bundles, medullary rays & pith. § Medullary rays: These are few layers of radially placed parenchymatous cells in between vascular bundles. § Vascular bundles: Large in number. They are arranged in a ring. Ring arrangement is a characteristic of dicot stem. Each vascular bundle is conjoint, open, and with endarch protoxylem. § Pith: Central portion of the stem. It has many rounded, parenchymatous cells with large intercellular spaces.
The vertical section of a dicot leaf through lamina shows 3 main parts: Epidermis, mesophyll & vascular system. o Epidermis: It covers both the upper surface (adaxial epidermis) and lower surface (abaxial epidermis) of the leaf. It has a conspicuous cuticle. Abaxial epidermis generally bears more stomata than the
adaxial epidermis. The latter may even lack stomata. o Mesophyll: The tissue between the upper and the lower epidermis. It is made up of parenchyma. They contain chloroplasts for photosynthesis. It has 2 types of cells: § Palisade parenchyma: It is adaxially placed. Made up of elongated cells arranged vertically and parallel to each other. § Spongy parenchyma: The oval or round and loosely arranged. It is situated below the palisade cells and extends to the lower epidermis. There are numerous large spaces and air cavities between these cells. o Vascular system: It includes vascular bundles. They can be seen in the veins and midrib. Size of vascular bundles is dependent on the size of the veins. The veins vary in thickness in the reticulate venation of dicot leaves. Vascular bundles are surrounded by a layer of thick walled bundle sheath cells.
The anatomy of monocot leaf is like that of the dicot leaf in many ways. It shows following differences: o Stomata are present on both surfaces of the epidermis. o Mesophyll is not differentiated into palisade and spongy parenchyma. o In grasses, certain adaxial epidermal cells along the veins modify themselves into large, empty, colourless cells. These are called bulliform cells. When the bulliform cells have absorbed water and are turgid, the leaf surface is exposed. When they are flaccid due to water stress, they make the leaves curl inwards to minimise water loss. o Parallel venation is reflected in the near similar sizes of vascular bundles (except in main veins).
Activity of the cambial ring
Secondary growth in a dicot stem – stages in transverse views
Spring wood and autumn wood
Heartwood and sapwood
of vascular cambium, the outer cortical & epidermis layers get broken. It is to be replaced to provide new protective cell layers. Hence another meristematic tissue called cork cambium (phellogen) develops, usually in the cortex.
Different stages of the secondary growth in a typical dicot root