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Chapter 1: The Scientific Method
Inductive Reasoning:
- Specific observations lead to generalizations
- Description based approach
- Particular → General
Scientific Process
- Observation: Orange is sweet
- Hypothesis: All oranges are sweet
- Prediction: If I taste all kinds of oranges, then they will all be sweet
- Test: Tasting all kind of oranges
- Travel to Madagascar oranges there are not sweet
- Hypothesis is falsified
Contract between science and knowledge:
- Initial skepticism on facts
- We ask honest questions, re-test what has been found
- Realism
- Acknowledge that the world is older and exists independently from my perception of it
- Realm of ideas < Real World
- Notions we have need to be realistic
- Rationality
- Deductive and inductive approach → Logic demonstrations from a scientist must be the result of
coherent steps
- Parsimony
- Methodological principle which states that acceptable theories are hypothetically the
most economical in assumptions
- Simplest option is usually true
- Methodological Materialism
- All that is experimentally accessible in the real world is material or has material origin
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Chapter 1: The Scientific Method

Inductive Reasoning:

  • Specific observations lead to generalizations
  • Description based approach
  • Particular → General

Scientific Process

  • Observation: Orange is sweet
  • Hypothesis: All oranges are sweet
  • Prediction: If I taste all kinds of oranges, then they will all be sweet
  • Test: Tasting all kind of oranges
    • Travel to Madagascar ⇒oranges there are not sweet
  • Hypothesis is falsified

Contract between science and knowledge:

  • Initial skepticism on facts
    • We ask honest questions, re-test what has been found
  • Realism
    • Acknowledge that the world is older and exists independently from my perception of it
    • Realm of ideas < Real World
    • Notions we have need to be realistic
  • Rationality
    • Deductive and inductive approach → Logic demonstrations from a scientist must be the result of coherent steps
    • Parsimony
      • Methodological principle which states that acceptable theories are hypothetically the most economical in assumptions - Simplest option is usually true
  • Methodological Materialism
    • All that is experimentally accessible in the real world is material or has material origin

Chapter 3: Charles Darwin and Evolution

Evolution and Natural Selection

  • Species resemble themselves because they share a common ancestor (not common environment)
    • Rejects fixity of species and accepts concept of descent with modifications (evolution)
    • Contradicts religious dogma of the time
    • Rejects Lamarck’s evolutionary mechanism (environmental determinism)
  • Finches are similar because they share a common ancestor → Finches of Galapagos Island
  • Rock Hyrax Example
  • Dugong Example
  • African Elephant
  • 99% of species that have lived on Earth are extinct
  • Most of the evolutionary branches finish in a dead-end (extinction) → Due to natural selection
  • Every human population has a tendency to increase geometrically, whereas the available resources to feed these populations increase arithmetically - Human population increases faster than its capacity to feed itself ⇒chaos (famine, sickness, war, etc.) & reduction in population size

Darwin’s First Observation:

  • All species can produce more offspring than their environment can sustain and many of these fail to survive and reproduce - Examples: - Mola mola (Ocean sunfish) - Maple tree and its samaras) - Puffball (spore cloud)

Darwin’s Second Observation

  • Members of a population often vary in their inherited traits
  • High variability in inherited traits
    • In our genes → transmitted to offspring
  • Examples
    • Amphidormus adamsii
    • Asian ladybird beetles
  • Natural Selection
    • Traits passed down that are favored while reproducing
    • Comes to the advantage of those being reproduced = reproductive success
    • Enables the emergence of adaptations

Important notions linked w/ natural selection:

  • Individuals don’t evolve, populations evolve
  • Only hereditary traits are subject to natural selection
  • For evolution to occur, we need genetic variability
  • Natural selection corresponds to differential reproductive success within a population from generation to generation
  • Natural selection enables individuals to become better adapted to their environment
  • Increased blood flow

Many regions in high altitudes were independently colonized by humans.

ANDES QINGHAI-TIBET

PLATEAU

  • Increase in hemoglobin concentration (RBC) - Increased blood flow
  • Natural selection keeps the individuals with the genetic traits that allow adaptation to high altitude
  • Individuals that have the mutation are at an advantage for survival and reproduction (production of offspring)

In three cases (moth, finch, humans) that hereditary traits that give a reproductive advantage to individuals in a population will be favoured

  • There will be a higher percentage of individuals carrying the adaptation
    • = natural selection
  • These adaptations can sometimes redefine a species and can occasionally define a new species

Chapter 4: Genetics, Neo-Darwinism, & Modern Synthesis

Modern additions to Mendel’s laws

Dominance

Incomplete Dominance:

  • Red is not TOTALLY dominant = mix
  • Example → Snapdragon

Incomplete Dominance

  • Example --> Palomino horse (Cc)

Codominance:

  • Example → Checkered chicken

Three concepts to use:

  1. Polygenism
    • When a phenotypic trait is under combined action of two or multiple genes (very common)
    • All characterized by continuous traits (height, skin color)
    • Eyes → Discrete; Skin color ⇒Parents
  2. Epistasis
    • When the effect of a gene hides or blocks the expression of another gene
  3. Pleiotropy
    • When a gene affects more than one trait
    • Manx cat →> M gene
    • Incomplete dominance compared to normal gene
    • Gene responsible for no tail = lethal (embryo reabsorbed) when homozygous ( ⇒death)
    • Heterozygous ⇒short tail (or no tail), only affects part of vertebral column

Chapter 5: Evolution of Populations

-images in notebook- -genotypic and phenotypic frequencies-

Hardy-Weinberg’s Principle

  • Population is in HW Equilibrium when:
    • Frequencies of alleles and genotype within a population will remain constant (according to p2+2pq+q2=1) from generation to generation as long as: - There are no mutations - Mating is done randomly - Size of the population is really large - There is no genetic (gene) flow (no migration of alleles between populations) - No natural selection
  • HW Principle describes a hypothetical population that does not evolve

Mutation

  • Changes in the DNA sequence of an organism
  • This is the source of genetic variability
  • Mutations are:
    • Random not directed
    • Transmissible (only in the gametes) to the next generation if it touches sex cells
    • Frequent throughout the gene pool but rare at each locus
    • Will influence allele frequencies but is a weak evolutionary force from generation to generation, espc in large population

Point Mutation

  • Additions, deletions, substitution of a base (A-C, instead of A-T)
  • Negative effect (Ehlers-Danlos Syndrome → fabrication of CT)
    • Linked with Dwarfism → Achondroplasia
  • Lethal effect (will cause death)
  • Found black guppies BUT died because they lost all genetic variability → deformed vertebral column, etc.

Choice of partners in relation to the phenotype (modifies HW)

Positive Assortative Mating Negative Assortative Mating

  • Autogamy or selfish of plants (pure lineage of Mendel) - Self-fertilization
  • Geographical proximity of individuals (population of mice in a barn)
  • In humans: mating according to height and skin color (not random)
  • Between individuals that look similar thus increasing homozygosity - Don’t look alike, and mate - Plants: Little change of self-fertilization, stamen and carpel = baby - Histocompatibility → female rejects own pollen therefore no SF - One species, two genders

Gene Flow

  • Migration: exchange of genes between populations
    • One species: gene flow is not possible with one population
  • Tends to standardize the genetic pool of the populations involved
    • Make it similar v quickly
    • Ex. Bass
    • Population ⇒Each lake does not interact with each other
    • Species ⇒Bass in general
  • Can play a similar role as mutations by introducing new genes in one of the populations

Example of Gene Flow → Parus major (Great Tit)

  • Mainland → Island migration
  • Central part survived, Eastern part was very different

Genetic Drift

  • Result of chance
  • Chance will have increasingly more impact on a population as the size of the population gets smaller
  • Small population
    • Genetic drift triggers decrease in genetic variability & decrease in heterozygosity
  • Large population
    • Cause little changes in allelic or genotypic frequency of a population
  • Larger the pop, less sampling error between expected and observed frequencies
  • If nothing affects allele frequencies, genetic drift will eventually result in the fixation of an allele and the elimination of all others for this locus
  • Probability that an allele will become fixed is equal to its frequency
  • Population bottleneck
    • Small populations → genetic drift = fixation of deleterious alleles and loss of genetic variability
    • This can correspond to an increase in rise of extinction
    • Cheetah → All of them are the same, when an epidemic occurs, most of them will die because all of them had the same alleles that were affected
    • Greater prairie chicken
      • Still some population left in the US
      • Average # of allele/locus

Selection for a continuously variable trait

Directional Selection

  • DDT and Mosquito Example
    • DDT → insecticide
    • MQ became resistant to DDT
    • High mortality, then very low mortality after 2 years
  • Medium Ground Finch example

Disruptive Selection

  • Birds and width of their lower jaw example → Pyrogentes ponceau
    • Measure width of lower jaw → correlated with seed they eat
    • Selection against average value
    • Intermediate size seed was not available
    • Extreme value is favored
    • Maybe important in speciation

Stabilizing Selection

  • Newborns that weighed 8lbs had a higher rate of survival than smaller or larger newborns
    • This stabilizing selection has diminished in many wealthy countries
  • All mammals have seven cervical vertebrae
    • Pleiotropic effect: one gene has many effects on the body
  • Not a standard habitat

Sexual Selection

  • Formulation of Darwin’s concept

Intersexual Selection

  • Individuals possess some specific hereditary traits are more susceptible than others to find partners
  • Ex. Widow bird
    • Do females birds prefer longer tails?
    • Females choose the male in this species
    • Very popular male → longer tails
  • Ex. Male stalk eyed flies
    • Longer eye stalk > shorter eye stalk
      • Healthier
  • Ex. Central American Birds
    • Moonwalk when they want to mate

Intrasexual Selection

  • Selection between individuals of same sex
    • Fight over who has sex
  • Confrontational behavior = ritualized combat
  • Ex. Kangaroos fighting

Penis Bone or Baculum → Walrus

  • Quick copulation
  • Also in dogs and cats
  • Facilitate penetration, lock penis inside their mate

Chapter 6: Adaptation

Evolutionary Compromise

  • Why are humans the only Primates choking on food?
    • Pay a price for something
  • Snout of a chimp (longer tongue) is longer than a humans
    • Reduces spaces between throat and tip of mouth
  • Back of mouth: Soft palate separates nasal area from mouth
    • Epiglottis: Cartilage shuts trachea

CHIMP HUMANS

  • Can swallow and the food is automatically blocked by epiglottis and trachea, there is not space for the food to go down the wrong pipe - Space between tip of palate and epiglottis - Swallow food and mechanism for epiglottis doesn’t work automatically - Space in between makes us vulnerable

Why only humans?

  • No a lot of space between palate
  • When can increase or decrease space, space between vocal cord is bigger in humans
  • Chimps can emit sound but not at the same capacity as humans
  • Humans tube allows for fast and clear pronunciation
    • Allows us to pronounce vowels clearly
  • Old adaptation

Studying Adaptations

  • Initial function of fins was to stabilize the fish body in its environment → adaptation
  • In terrestrial animals (tetrapods) the limbs (that evolved from fins) are used to support the body in an aerial environment - Limbs are exaptations for terrestrial mobility (locomotion)
  • Exaptation allows us to have a historical outlook to adaptation
    • Current function is not what it initially was Ex. Hair
  • Adaptation → thermoregulation
  • Exaptation → to dissipate heat for elephants
  • They have thinner skin in some areas
  • Trunk, ears, these random patches have hot spots
  • Origin of hair was not to preserve heat, it was to get rid of it, but NOW it preserves heat

Ex. Feathers

  • Preservation of heat
  • Dinosaurs were not endotherms, they accumulated hear with feathers
  • Origin is not known, but it’s for sure that it wasn’t for flight, possibly sexual selection

Chapter 7: Geographic Variation and Speciation

Evolution of the loss of fur and skin color in humans

  • 6-8 million years ago, the hominins (all human species) shared a common ancestor with the chimpanzee (pale skin covered of a dark fur)
  • Sahelanthropus tchadensis → oldest hominin known

A. afarensis

  • Lived in tropical forests
    • A lot of hair
  • Bipedal, and arboreal
    • Long arms, feet with capacity to climb attachment of VC, under the cranium
  • Diet of fruits, tubers, and leaves
    • Frugivores
  • Sedentary way of life
    • Limited movement
  • Australopithecus afarensis → 3.7-3 Myr
  • Skin of all hominis species (including Homo sapiens ) was dark
  • Hypothesis:
  • Find an inherited trait that gave those who had it a positive survival and reproduction differential (adaptation) from generation → generation compared to other members of the population
  • Fur
  • Protection from abrasion to skin
  • Hair reacts to cold and strong emotion
  • Loss of hair = facial expressions became important
  • Dark skin
  • For over 1 Myr
  • The skin of ALL hominins species was dark
  • Including homo sapiens

Vitamin D and Pale Skin 3 

  • Synthesized in skin by UV rays
  • Helps absorption of Ca 2+^ in the gut
    • Deficiency of Calcium ⇒rickets, other diseases
  • Direct impact on reproductive success of affected individuals
  • Pale skin in low UV radiation → maximize absorption of UV rays and the survival of the human population in areas of UV deficiency (adaptation)
  • More North = paler skin
  • Migration = 100 000 years ago
    • Humans invade northern habitats more recently →. 40 000 years ago
    • Evolve paler skin to maximize UV absorption in areas where UV radiation is weak for adequate synthesis of Vitamin D by the skin 3 
  • Humans can compensate for this (10-15 000 years ago) through diet (hunting, fishing, domestication)

Conclusion

  • Loss of fur in hominins linked to lifestyle
    • Due to climate change → 1.2 Myr
  • Skin = dark after loss of fur → 1 Myr
  • Colonization up north = paler skin → maximize D absorption 3 
    • Dark skin = synthesize D at a slower rate 3 
  • Selective advantage of darker skin on high UV intensity areas = minimize degradation of folic acid by UV rays

Color of Skin

  • Variable polygenic trait, explains pigment intensity variability
  • Adaptation to UV radiation
  • Indicator of environment in which population has lived

Origin of Species

  • Eastern Meadowlark ( Sturnella Magna) and Western Meadowlark ( Sturnella

neglecta )

  • Minimal phenotypic differences (except for their song and behavior that are very distinct)

Sympatric speciation:

  • New species appears within population (speciation without geographic isolation):
  • Polyploidy (30-40% of plants): multiplying the normal number of chromosomes
  • Can happen when the chromosomes do not separate during meiosis → produces diploid games instead of haploids
  • Rare in animals
  • Ex. Apple maggot v. Thornapple maggot
  • Two morphs of Rhagoletis pomonella in the process of sympatric speciation

Hybrid Zone

  • Ex. Toads
    • Yellow bellied toad → Bombina variegata
      • Lives at higher altitudes
    • Fire bellied toad → Bombina bombina
      • Lives at lower altitudes
    • Both came to the middle altitude or the hybridzone and more of them survived there than in either of the altitudes
  • Ex. Grolar bear
    • Rupture of reproductive isolation between grizzly and polar bear
    • Exception? Consequence of global warming?
  • Enhancement of reproductive barriers in the contact area (hybrid zone)
    • Character displacement
      • Differences between two related species are often greater in sympatric areas
        • 2 species of Galapagos finches
  • Enhancement of reproductive barriers in flycatchers
  • 3500 Myr since LUCA
  • Human species is the only one terminal twig in the bush of life
  • Luca is at the center of things, with 3 branches of life around it
  • Some branches will stop meaning they dies (extinction)
  • All species share a common ancestor that is at the center of the bush
  • WE ARE ALL THE SAME → we make have a different way of life, but we are all equal in terms of evolution of species because we all share 3500 million years of history

Homology and Homoplasy

Homologous structures

  • Indicative of a common ancestor and the data that is used to reconnect phylogenies
  • Ex. Humerus in a human vs humerus in cat, whale, and bat
  • Limb of species have different functions and can be found in different species, but they are homologous structures that indicate a common ancestor

Homoplasy

NO HOMOPLASIES IN PHYLOGENY

  • Convergent Evolution
    • Ability to hover in sugar gliders v flying squirrels → might be related in terms of phylogeny but they might have evolved the gliding trait independently
    • Convergent evolution is a problem when you describe phylogeny because not all characters that look alike have common ancestors
  • Evolutionary Reversion
    • Absence of fur in whales

Taxonomy and Classification

  • Biologists use binomial system to identify living organisms
  • Homo = name of genus
    • Signifies “man”
  • Sapiens = name of species
    • Signifies: intelligent, wise and reasonable
  • Classification
    • System of words used to group species into increasingly generalized categories
    • Hierarchical system
      • Sequential and orderly arrangement of categories
  • Taxonomic rank → taxon
  • The classification needs to reflect on the phylogeny

Phylogenetics

  • Tree with dichotomous branches (cladogram)
  • Bifurcation point (branching point)
    • Common ancestor denotes related groups
  • Cladogram indicates the sequence of appearance of taxa
  • No time frame incorporated in cladogram
  • Classification reflects ramifications of the cladogram, homologous characters help define the order of sequence

Branching point and polytomy

Taxon D, E, F → Unresolved Basal Taxon: defines relationship in A-F

  • Basis to compare evolution within our group

Cladistics:

  • Taxa should be based on shared derived characters aka
    • Synapomorphy
    • Homology
      • Material upon we build classification
  • Not shared ancestral characters aka
    • Symplesiomorphy
      • Do not want this!
      • A character might be either derived or ancestral character → distinguish this!
  • Only valid group
    • Monophyletic group

Monophyletic groups → contains all and only the descendants from a common ancestor NON MONOPHYLETIC GROUPS: (WE DON’T WANT THIS) Paraphyletic groups → does not contain all the descendants from a common ancestor Polyphyletic groups → contains descendants from several ancestors

  • Only way to find an answer is to look at birds and crocodilians → birds came from dinosaurs and crocodiles came before dinosaurs
  • Hypothesis based on cladogram: birds and crocs take care of their young → if oldest and most derived groups related to dinos take care of their young, it's safe to assume that based on historical data that dinos took care of their young too

Chapter 9: Biosphere and Ecology

Ecology: Definitions, Concepts, and Issues

Ecology

  • Scientific study of interactions between organisms and between organisms and their environment
  • Environment:
    • Abiotic Factors → interactions between organism and environment
    • Biotic Factors → interactions between organisms

Scope of Ecology

  • Organism ecology
    • Interactions between organism and environment
      • Physiology, behavior
      • Main example = evolution
  • Population ecology
    • Groups of organisms of one species living in the same age (dynamics of population size fluctuations)
  • Community ecology
    • Groups of populations of different species in an area
      • How interactions between species (predation, competition) affect the structure and organization of communities
  • Ecosystem ecology
    • Community of organisms in an area and the interaction of organisms with physical factors (study of energy flow, chemical cycling)
    • How animals interact with their environment
    • ecosystem approach is more important and prioritize ecosystems with endangered species
  • Landscape ecology
    • Mosaic of connected ecosystems (study of factors controlling of energy, materials and organisms across multiple ecosystems)
    • Ex: Ottawa has a green belt, conscious attempt to keep a belt of forest around the city, important for fauna and flora (helps maintain), have corridors to connect ecosystems
  • Global ecology
    • Study of the biosphere
    • Sum of all ecosystems (study of regional exchange of energy and materials its global impact on the biosphere)
    • Activity done in one part of the ecosystem and affects the entire planet ( global warming)

Precautionary Principles

  • Strategy to cope with possible risks where scientific understanding is yet incomplete
    • Better to be safe than sorry
  • Earth’s climate is warming up due to human activity
  • Interaction between society and ecology
  • Always uncertainty with science doesn't matter what kind of science
  • Scientists might give warning → ex: climate change, scientific evidence indicated that ifwe keep emitting the amount of carbon we are, temperature will rise
  • We have to find a way to come with it ( summits that are put together between countries)
  • It's going to cost more to repair things than prevent it

Ex: lack of cod in the Maritimes, there was a limited supply due to all the fishing done. Government warned that if people didn’t stop fishing the cod, there would be none left à ultimately no one took the precaution seriously, overfished cod, Maritimes fishing business collapsed

MAYBE if precautionary principle was applied, we might still have cod

Climate

Abiotic Factors

  • Light, Temperature, Wind, Precipitation
  • In polar regions:
    • Solar rays are slanted whereas they are vertical in equatorial regions
    • Solar energy spreads over a large surface area
    • Temperature varies depending on where you are on the planet
  • Earth is tilted on its axis at 23.5 degrees angle
    • Extratropical regions have seasons
    • Tropical regions have little differences between seasons and days
    • Angles of the earth has an effect on our climate à makes seasons because the fact of what part of the earth is closer or farther away from the sun determines the weather
    • If the earth became flat, we would lose our seasons
    • Angles also determine how much light we get (winter and summer equinox)
    • All facts are impact on species

Air circulation and precipitations at a global scale

  • Winds are made with friction between atmosphere and planet → the friction is the cause of win
  • Wind is not straight
  • Descending dry air absorbs moisture
  • Ascending moist air releases moisture
  • Circulation of surface water in the oceans influence climate
  • All currents are linked
  • Combination between warm and cold currents
  • Currents have an impact on climate on land
  • Effect of large bodies of water on the climate
    • Cool air comes from the ocean
    • Land absorbs and emits more heat making the air warm